7 Pacific islands with tortoises visited by Darwin in 1835 (9)
8 Resort city in western Florida (5)
10 Desert in south-western Africa (8)
11 Belgian seaport with ferry links to Dover and Folkestone (6)
12 Irish Free State (4)
13 Republic in eastern Africa (8)
15 Mexican state, capital Culiacán (7)
17 Spanish province and city in Extremadura (7)
20 Cumbrian city (8)
22 The ´Eternal City` (4)
25 Ancient Roman city, scene of Hannibal's victory in 216 B.C. (6)
26 London's third airport (8)
27 Spanish city and naval base, Drake singed the King of Spain's beard here in 1587 (5)
28 Any two places or regions on diametrically opposite sides of the Earth (9)

1 Country, capital Tokyo (5)
2 African state, formerly Nyasaland (6)
3 Location, on the 28th of June 1914 of the assassination of Archduke Franz Ferdinand (8)
4 Ancient city in Greece (7)
5 Ancient city near Tunis (8)
6 Mountain range extending the length of the Italian peninsula (9)
9 Seat of W German government 1949-90 (4)
14 Mountain range containing Everest (9)
16 "His glowing axle doth allay in the steep --- stream" [Milton] (8)
18 Arm of the Mediterranean between Yugoslavia and Albania on the east and Italy on the west (8)
19 A port city in northeastern Sicily (7)
21 Canal connecting the Mediterranean Sea and the Indian Ocean (4)
23 Capital of the Russian Federation (6)
24 City in southwestern Spain that is famous for making sherry (5)